<inline-graphic xlink:href="nl-49-017-i001.jpg" xlink:type="simple" id="oo_1568715.jpg"/> On January 16th, our dear colleague and distinguished entomologist Enrique García-Barros Saura passed away at the age of 66. All of us who knew him, and especially those of us who cultivated his friendship, hold an indelible memory of his worth, his scientific and intellectual excellence, and above all, his humanity. We never wished to write this obituary, the words of which we still struggle to comprehend and accept, and we dare to publish it only as a tribute to his memory and to his person. Enrique was born in Madrid on June 11th, 1959, into a middle-class family. His father, Dr. Enrique García-Barros Bernabéu, was a psychiatrist; his mother, María Pilar Saura Atarés, was the sister of the artists Antonio (a painter) and Carlos (a filmmaker) Saura. The eldest of three siblings, he was a very curious, observant, and introverted child who loved animals. Enrique recounted how, as a child, he had won a photography contest with a portrait of his tortoise, Cleopatra, and that he always had pets: hamsters, lizards, fishes... He often heard the monkeys, bears, and other animals at the zoo in the Retiro, a park in the centre of Madrid, that was right across the street from his childhood home. At the age of four, he started kindergarten (as it was called then) at the “Santa María del Pilar” school in the Retiro neighbourhood, also very close to his home, and he finished his secondary education and university access exams there. In October 1976 and until December 1981, he studied for a Biology Degree at the Universidad Autónoma de Madrid (<abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>). Some of Enrique’s earliest memories are fondly recalled by one of the signatories of this note (JLV), who was his Zoology professor. Perhaps the oldest of these memories refers to the collection of the 100 animals that, at that time, students were required to deliver. Most of the specimens were of common insects collected around the buildings that housed the laboratories on the campus at Cantoblanco; there were also shells of molluscs found on the beaches (and perhaps in amongst the paella!), earthworms preserved in alcohol, lizards killed at the roadside, etc. García-Barros as a student included among his specimens a paper butterfly that he had painted himself (a <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Thymelicus">Thymelicus</tp:taxon-name-part></tp:taxon-name></italic>, to be precise) which faithfully imitated a real specimen, and, of course, although it was perfectly pinned and labelled, it was discovered. As an excuse, and after the resultant telling off, Enrique argued that there was no reason to kill the butterfly, and that a model was more than enough. It must have been the first lesson that he gave us. On June 16th, 1982, he defended his master thesis, entitled “The Butterflies of La Alcarria,” which was awarded the highest distinction. Shortly afterwards, he began his PhD thesis under the guidance of Drs. Joaquín Templado and José Luis Viejo. Given the experience gained from his undergraduate thesis, and at Dr. Templado’s suggestion, Enrique undertook a comparative biological study of four butterfly species, scarcely known at that time. That thesis, entitled “Comparative Biological Study of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">Hipparchia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="semele">semele</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">H.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="statilinus">statilinus</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">H.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="alcyone">alcyone</tp:taxon-name-part></tp:taxon-name></italic>, and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Pseudotergumia">Pseudotergumia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="fidia">fidia</tp:taxon-name-part></tp:taxon-name></italic> (Lep. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Satyridae">Satyridae</tp:taxon-name-part></tp:taxon-name>),” was defended at the Universidad Autónoma de Madrid on July 1, 1987, and unanimously awarded the highest distinction, “<italic>cum laude</italic>.” It was the first entomology thesis completed entirely within the <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>’s Department of Biology. In October 1984, he obtained a position as an assistant teacher in the Department of Biology at the <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>’s Faculty of Sciences. Within this category, he completed a postdoctoral fellowship at the Zoologiska Institutionen, Stockholm University, from March 1990, to April 1991, under the supervision of Professor Christer Wiklund, one of the leading figures in the study of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name> ecology in Europe. His time in Sweden provided a significant boost to his scientific career, as he began collaborating with prominent European entomologists, particularly with his mentor in Stockholm, Dr. Wiklund. Taken together, the work of García-Barros and Wiklund (1992–1997) introduced a comparative and functional approach to European lepidopterology, connecting developmental physiology, ecology, and evolution, and made up a direct step towards Enrique García-Barros’s subsequent studies on allometry, body size, and macroecological patterns, which would eventually be integrated into spatial and cartographic analyses of biodiversity. In July 1993, he gained the position of Professor of Zoology in the Department of Biology at the Universidad Autónoma de Madrid and began to take on greater teaching responsibilities. In the subsequent years, he taught theoretical and practical courses in the Biology Degree program, particularly the subject of Zoology in the initial years of this degree. He also collaborated in teaching the Applied Entomology course in the Master’s program in Biodiversity and some other subjects. In recent years, he taught courses in the Biology degree program and the Biodiversity and Ecology master’s programs at the Universidad Autónoma de Madrid, including Zoology (Biology degree), Advanced Laboratory (Taxonomy Laboratory, 3rd year, Biology degree), Biodiversity of Terrestrial Organisms (Master in Biodiversity, <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>), and Biogeography (Master’s degree in Ecology, <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>-UCM). He also supervised numerous undergraduate and Master’s theses. A testament to his tireless dedication, open and innovative spirit, and originality was the fact that he was supervising, together with one of the signatories (JLV), an undergraduate thesis in the Environmental Sciences degree entitled “Environmental aspects of the work of Hieronymus Bosch.” Furthermore, he coordinated the two published editions of the university’s “Manual de Prácticas de Zoología”, which compile the practical guides for this subject and is widely used by students. He is a co-author of several of these guides that included original drawings by himself. His research focused primarily on the Order <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>, and in particular on Iberian butterflies (superfamily <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name>). He had been interested in the chorology, biogeography, phylogeny, behaviour, morphology, ecology, biology, and conservation of these insects. Of Dr. García-Barros’s 169 publications, it is extremely difficult to single out any one in particular, but if we had to, we would say it was his direction, co-authorship, and promotion of “Fauna iberica” volume 37, dedicated to butterflies: Fauna iberica. Vol. 37. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name>. García-Barros, E., Munguira, M. L., Stefanescu, C. & Vives Moreno, A. (2013), 1216 pages. This book constitutes the most comprehensive taxonomic and biogeographical synthesis on the butterflies of the Iberian Peninsula and Balearic Islands. It systematically integrates morphological diagnoses, biological, ecological, and distributional data, supported by a documentary and faunal database with records for over more than a century. From a historical perspective, it represents the culmination of the Iberian faunal tradition, while also incorporating modern criteria of chorology, conservation status, and geographic variability, making it an essential reference work for research, biological mapping, and the conservation of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> in Spain. Among the publications for which he was the leading author, two stand out: the “Atlas of the Butterflies of the Iberian Peninsula and Balearic Islands (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> & <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Hesperioidea">Hesperioidea</tp:taxon-name-part></tp:taxon-name>),” (2004), which has been widely cited and has served as the basis for dozens of collaborative works with numerous authors; and “Parsimony Analysis of Endemicity and Its Application to Animal and Plant Geographic Distributions in the Ibero-Balearic Region (Western Mediterranean),” published in the Journal of Biogeography in 2002, which is his most cited work due to its remarkable originality. Another noteworthy work was the Atlas and Red Book of Spanish Butterflies, for which he was a member of the coordination team, along with two of this note authors (HR, MLM). Sadly, Enrique did not live to see this important contribution published, as he passed away three months after its submission. Enrique’s attendance and presentations at national and international conferences were a constant since 1982, when he attended the “V Jornadas de la Asociación española de Entomología” held in Valencia with his presentation on the “Butterfly Fauna (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Hesperioidea">Hesperioidea</tp:taxon-name-part></tp:taxon-name>) of the Alcarria Region.” He began his participation in international conferences in 1986, attending the “5th European Congress of Lepidopterology of the Society for European Lepidopterology” (<abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev>) held in Hungary, with the presentation “Period of pre-oviposition and notes on the biology of the Grayling, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">Hipparchia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="semele">semele</tp:taxon-name-part></tp:taxon-name></italic> (L.) in Central Spain.” Subsequently, Enrique attended many more <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> conferences; his expertise, regular participation, and fluent English made him one of the most recognised European lepidopterists, and he also served on the Council of this prominent scientific society. For over 30 years, he participated in the <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> biennial congresses at numerous European venues. He presented more than 90 papers at the various congresses in which he participated. It is also worth mentioning his contribution as one of the organisers of the “X European Congress of Lepidopterology”, held in Miraflores de la Sierra (Madrid), May, 1996. In addition to attending the congresses of <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> and the “Asociación Española de Entomología” (both national and Iberian), he has presented works at other conferences of societies such as the “Sociedad Hispano-Luso-Americana de Lepidopterología” (<abbrev xlink:title="Sociedad Hispano-Luso-Americana de Lepidopterología">SHILAP</abbrev>), the “Real Sociedad Española de Historia Natural”, the “Institute of Ecology and Systematics”, and at international congresses such as “Future of Butterflies in Europe”, the “International Congress of Entomology, Biology of Butterflies, Behavioural Ecology, Butterfly Ecology and Evolution”, and those organized by the “International Biogeography Society.” This demonstrates his interest in the biology, evolution, behaviour, biogeography, and conservation of butterflies. A comprehensive list of his publications can be found in the Appendix. He also co-directed two doctoral theses with Miguel L. Munguira. The first, defended in 2003 by Patrícia Garcia-Pereira, focused on “Butterflies of Continental Portugal: Faunistics, Biogeography, and Conservation.” This thesis assessed the knowledge of Portugal’s lepidopterological fauna, identifying areas with the greatest species richness and diversity and proposing appropriate conservation strategies. The second, defended in 2008 by Helena Romo Benito, was entitled “Geographical Diversity of Ibero-Balearic Butterflies.” This work provided an analysis of the distribution and richness of the 226 species of butterflies in the Ibero-Balearic region. It was the first time that butterflies were used to delimit biogeographical regions based on the species they harbour. This thesis also estimates butterfly sampling effort and associated biases, evaluate the national coverage of the network of protected areas for these insects, and develop initial models of potential distribution for rare or threatened species. Enrique also co-directed, together with Ana Luz Márquez and Raimundo Real from the Universidad de Málaga (<abbrev xlink:title="Universidad de Málaga">UMA</abbrev>), the thesis defended in 2022 by Antonio Pulido entitled “Application of fuzzy logic to the analysis of the spatial structure of diurnal butterfly populations in the Iberian Peninsula”. As a field entomologist, his tenacity was noteworthy. To find the elusive larvae of the satyrids he studied, he would amble through clumps of grass with an enormous comb and collect the caterpillars in a metal dish. His tremendous effort to complete the butterfly data for Spanish areas lacking records was also remarkable. He took advantage of both personal and university trips to stop over in unexplored regions. The butterfly database used for the two editions of the “Atlas de Mariposas Españolas” compiles his numerous field observations, scattered throughout Spain. With endless creativity, Enrique got surprising and often brilliant proposals and initiatives. He made excellent sketches and possessed a capacity for abstraction and synthesis far superior to most of us mere mortals, making him such a pleasure to work with. He constantly surprised us with ingenious ideas, collages, humorous posters, and those small details that always brought a smile to our faces. He was not one to delegate, but he loved helping his colleagues and family. His passion was research, without neglecting his teaching; his classes were engaging and distinctive, designed to cultivate critical thinking skills in students who questioned everything. As a colleague, Enrique García-Barros was an exceptional person. He always exuded an optimism, tinged with skepticism and irony. His idiosyncrasies included his humming and his soft whistling in the department corridors which were legendary, as was his intelligent humour with colleagues and students. It was a pleasure to work with him; he was always willing, attentive, a good listener, and generous. Discreet and modest, even somewhat introverted, he rarely expressed his worries, which, like anyone else, he surely had. He faced his short and fatal illness with the same tranquillity as the rest of his life. His family, friends, colleagues, and all who knew him will miss him utterly. Rest in peace, Enrique. <bold>Y ME DECÍS LUEGO</bold> Se marcha Enrique, El pasillo queda silente, Sin el silbo, sin el canturreo, Quizá un lejano rumor de voces inconcretas. Luego Se apaga la luz y todos lloran, Pero enseguida se encienden candiles y bengalas; Son chispas, fogonazos, flashes, El fondo sigue siendo oscuro y silencioso Pero hay luces fugaces Que alegran las miradas, Ecos lejanos, Pero perceptibles, Que recuerdan que no se fue del todo, Que permanece tanto En livianos garabatos, Como en dibujos olvidados Y, cómo no, en sus escritos, Sólidos, magníficos, creativos, De portentosa imaginación Y geniales ocurrencias. No, no se fue del todo, Escuchad esos ecos, Recordad esas vivencias Y me decís luego. <bold>AND TELL ME LATER</bold> Enrique leaves, The hallway falls silent, Without the whistling, without the humming, Perhaps a distant murmur of indistinct voices. Then The light goes out and everyone cries, But soon candles and flares are lit; They are sparks, flashes, bursts, The background remains dark and silent But there are fleeting lights That brighten our gazes, Distant echoes, But perceptible, That remind us he didn’t leave completely, That he remains both In light scribbles, As in forgotten drawings And, of course, in his writings, Solid, magnificent, creative, Of prodigious imagination And brilliant insights. No, he didn’t leave completely, Listen to those echoes, Remember those experiences And tell me later. </sec> </body> <back> <app-group> <app id="app1"> <title>Appendix 1. List of publications

urn:lsid:arphahub.com:pub:E0185C18-FE79-5ADE-9877-ED333312DD4F

urn:lsid:zoobank.org:pub:EF082B8D-8FD0-41B8-BC24-3D1190FEC17F

Nota Lepidopterologica

0342-7536 2367-5365

Pensoft Publishers

10.3897/nl.49.189708

189708

In Memoriam

Lepidoptera

Cenozoic

Europe

Dr. Enrique García-Barros Saura. In Memoriam

Viejo Montesinos

José Luis

joseluis.viejo@uam.es https://orcid.org/0000-0003-3160-6892 1 Romo Benito

Helena

https://orcid.org/0000-0002-5729-1342 1 Munguira

Miguel López

https://orcid.org/0000-0002-1554-4380 1

Departamento de Biología, Área de Zoología, Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Facultad de Ciencias, Universidad Autónoma de Madrid, Madrid, Spain; joseluis.viejo@uam.es, helena.romo@uam.es, l.munguira@hotmail.com

Universidad Autónoma de Madrid

Madrid

Spain https://ror.org/01cby8j38

Subject Editor: Carlos Lopez-Vaamonde

2026

25 03 2026

49 17 33 AA61DDA5-651C-5765-BBD7-E54C46191A54 83CB262B-18EF-4DAA-9874-329F7A4E9FE4 24 02 2026 28 02 2026

José Luis Viejo Montesinos, Helena Romo Benito, Miguel López Munguira

This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

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<inline-graphic xlink:href="nl-49-017-i001.jpg" xlink:type="simple" id="oo_1568715.jpg"/> On January 16th, our dear colleague and distinguished entomologist Enrique García-Barros Saura passed away at the age of 66. All of us who knew him, and especially those of us who cultivated his friendship, hold an indelible memory of his worth, his scientific and intellectual excellence, and above all, his humanity. We never wished to write this obituary, the words of which we still struggle to comprehend and accept, and we dare to publish it only as a tribute to his memory and to his person. Enrique was born in Madrid on June 11th, 1959, into a middle-class family. His father, Dr. Enrique García-Barros Bernabéu, was a psychiatrist; his mother, María Pilar Saura Atarés, was the sister of the artists Antonio (a painter) and Carlos (a filmmaker) Saura. The eldest of three siblings, he was a very curious, observant, and introverted child who loved animals. Enrique recounted how, as a child, he had won a photography contest with a portrait of his tortoise, Cleopatra, and that he always had pets: hamsters, lizards, fishes... He often heard the monkeys, bears, and other animals at the zoo in the Retiro, a park in the centre of Madrid, that was right across the street from his childhood home. At the age of four, he started kindergarten (as it was called then) at the “Santa María del Pilar” school in the Retiro neighbourhood, also very close to his home, and he finished his secondary education and university access exams there. In October 1976 and until December 1981, he studied for a Biology Degree at the Universidad Autónoma de Madrid (<abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>). Some of Enrique’s earliest memories are fondly recalled by one of the signatories of this note (JLV), who was his Zoology professor. Perhaps the oldest of these memories refers to the collection of the 100 animals that, at that time, students were required to deliver. Most of the specimens were of common insects collected around the buildings that housed the laboratories on the campus at Cantoblanco; there were also shells of molluscs found on the beaches (and perhaps in amongst the paella!), earthworms preserved in alcohol, lizards killed at the roadside, etc. García-Barros as a student included among his specimens a paper butterfly that he had painted himself (a <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Thymelicus">Thymelicus</tp:taxon-name-part></tp:taxon-name></italic>, to be precise) which faithfully imitated a real specimen, and, of course, although it was perfectly pinned and labelled, it was discovered. As an excuse, and after the resultant telling off, Enrique argued that there was no reason to kill the butterfly, and that a model was more than enough. It must have been the first lesson that he gave us. On June 16th, 1982, he defended his master thesis, entitled “The Butterflies of La Alcarria,” which was awarded the highest distinction. Shortly afterwards, he began his PhD thesis under the guidance of Drs. Joaquín Templado and José Luis Viejo. Given the experience gained from his undergraduate thesis, and at Dr. Templado’s suggestion, Enrique undertook a comparative biological study of four butterfly species, scarcely known at that time. That thesis, entitled “Comparative Biological Study of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">Hipparchia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="semele">semele</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">H.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="statilinus">statilinus</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">H.</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="alcyone">alcyone</tp:taxon-name-part></tp:taxon-name></italic>, and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Pseudotergumia">Pseudotergumia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="fidia">fidia</tp:taxon-name-part></tp:taxon-name></italic> (Lep. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family" reg="Satyridae">Satyridae</tp:taxon-name-part></tp:taxon-name>),” was defended at the Universidad Autónoma de Madrid on July 1, 1987, and unanimously awarded the highest distinction, “<italic>cum laude</italic>.” It was the first entomology thesis completed entirely within the <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>’s Department of Biology. In October 1984, he obtained a position as an assistant teacher in the Department of Biology at the <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>’s Faculty of Sciences. Within this category, he completed a postdoctoral fellowship at the Zoologiska Institutionen, Stockholm University, from March 1990, to April 1991, under the supervision of Professor Christer Wiklund, one of the leading figures in the study of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name> ecology in Europe. His time in Sweden provided a significant boost to his scientific career, as he began collaborating with prominent European entomologists, particularly with his mentor in Stockholm, Dr. Wiklund. Taken together, the work of García-Barros and Wiklund (1992–1997) introduced a comparative and functional approach to European lepidopterology, connecting developmental physiology, ecology, and evolution, and made up a direct step towards Enrique García-Barros’s subsequent studies on allometry, body size, and macroecological patterns, which would eventually be integrated into spatial and cartographic analyses of biodiversity. In July 1993, he gained the position of Professor of Zoology in the Department of Biology at the Universidad Autónoma de Madrid and began to take on greater teaching responsibilities. In the subsequent years, he taught theoretical and practical courses in the Biology Degree program, particularly the subject of Zoology in the initial years of this degree. He also collaborated in teaching the Applied Entomology course in the Master’s program in Biodiversity and some other subjects. In recent years, he taught courses in the Biology degree program and the Biodiversity and Ecology master’s programs at the Universidad Autónoma de Madrid, including Zoology (Biology degree), Advanced Laboratory (Taxonomy Laboratory, 3rd year, Biology degree), Biodiversity of Terrestrial Organisms (Master in Biodiversity, <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>), and Biogeography (Master’s degree in Ecology, <abbrev content-type="institution" xlink:title="Universidad Autónoma de Madrid">UAM</abbrev>-UCM). He also supervised numerous undergraduate and Master’s theses. A testament to his tireless dedication, open and innovative spirit, and originality was the fact that he was supervising, together with one of the signatories (JLV), an undergraduate thesis in the Environmental Sciences degree entitled “Environmental aspects of the work of Hieronymus Bosch.” Furthermore, he coordinated the two published editions of the university’s “Manual de Prácticas de Zoología”, which compile the practical guides for this subject and is widely used by students. He is a co-author of several of these guides that included original drawings by himself. His research focused primarily on the Order <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>, and in particular on Iberian butterflies (superfamily <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name>). He had been interested in the chorology, biogeography, phylogeny, behaviour, morphology, ecology, biology, and conservation of these insects. Of Dr. García-Barros’s 169 publications, it is extremely difficult to single out any one in particular, but if we had to, we would say it was his direction, co-authorship, and promotion of “Fauna iberica” volume 37, dedicated to butterflies: Fauna iberica. Vol. 37. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name>. García-Barros, E., Munguira, M. L., Stefanescu, C. & Vives Moreno, A. (2013), 1216 pages. This book constitutes the most comprehensive taxonomic and biogeographical synthesis on the butterflies of the Iberian Peninsula and Balearic Islands. It systematically integrates morphological diagnoses, biological, ecological, and distributional data, supported by a documentary and faunal database with records for over more than a century. From a historical perspective, it represents the culmination of the Iberian faunal tradition, while also incorporating modern criteria of chorology, conservation status, and geographic variability, making it an essential reference work for research, biological mapping, and the conservation of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> in Spain. Among the publications for which he was the leading author, two stand out: the “Atlas of the Butterflies of the Iberian Peninsula and Balearic Islands (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order" reg="Lepidoptera">Lepidoptera</tp:taxon-name-part></tp:taxon-name>: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> & <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Hesperioidea">Hesperioidea</tp:taxon-name-part></tp:taxon-name>),” (2004), which has been widely cited and has served as the basis for dozens of collaborative works with numerous authors; and “Parsimony Analysis of Endemicity and Its Application to Animal and Plant Geographic Distributions in the Ibero-Balearic Region (Western Mediterranean),” published in the Journal of Biogeography in 2002, which is his most cited work due to its remarkable originality. Another noteworthy work was the Atlas and Red Book of Spanish Butterflies, for which he was a member of the coordination team, along with two of this note authors (HR, MLM). Sadly, Enrique did not live to see this important contribution published, as he passed away three months after its submission. Enrique’s attendance and presentations at national and international conferences were a constant since 1982, when he attended the “V Jornadas de la Asociación española de Entomología” held in Valencia with his presentation on the “Butterfly Fauna (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Papilionoidea">Papilionoidea</tp:taxon-name-part></tp:taxon-name> and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="superfamily" reg="Hesperioidea">Hesperioidea</tp:taxon-name-part></tp:taxon-name>) of the Alcarria Region.” He began his participation in international conferences in 1986, attending the “5th European Congress of Lepidopterology of the Society for European Lepidopterology” (<abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev>) held in Hungary, with the presentation “Period of pre-oviposition and notes on the biology of the Grayling, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hipparchia">Hipparchia</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="semele">semele</tp:taxon-name-part></tp:taxon-name></italic> (L.) in Central Spain.” Subsequently, Enrique attended many more <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> conferences; his expertise, regular participation, and fluent English made him one of the most recognised European lepidopterists, and he also served on the Council of this prominent scientific society. For over 30 years, he participated in the <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> biennial congresses at numerous European venues. He presented more than 90 papers at the various congresses in which he participated. It is also worth mentioning his contribution as one of the organisers of the “X European Congress of Lepidopterology”, held in Miraflores de la Sierra (Madrid), May, 1996. In addition to attending the congresses of <abbrev xlink:title="Society for European Lepidopterology">SEL</abbrev> and the “Asociación Española de Entomología” (both national and Iberian), he has presented works at other conferences of societies such as the “Sociedad Hispano-Luso-Americana de Lepidopterología” (<abbrev xlink:title="Sociedad Hispano-Luso-Americana de Lepidopterología">SHILAP</abbrev>), the “Real Sociedad Española de Historia Natural”, the “Institute of Ecology and Systematics”, and at international congresses such as “Future of Butterflies in Europe”, the “International Congress of Entomology, Biology of Butterflies, Behavioural Ecology, Butterfly Ecology and Evolution”, and those organized by the “International Biogeography Society.” This demonstrates his interest in the biology, evolution, behaviour, biogeography, and conservation of butterflies. A comprehensive list of his publications can be found in the Appendix. He also co-directed two doctoral theses with Miguel L. Munguira. The first, defended in 2003 by Patrícia Garcia-Pereira, focused on “Butterflies of Continental Portugal: Faunistics, Biogeography, and Conservation.” This thesis assessed the knowledge of Portugal’s lepidopterological fauna, identifying areas with the greatest species richness and diversity and proposing appropriate conservation strategies. The second, defended in 2008 by Helena Romo Benito, was entitled “Geographical Diversity of Ibero-Balearic Butterflies.” This work provided an analysis of the distribution and richness of the 226 species of butterflies in the Ibero-Balearic region. It was the first time that butterflies were used to delimit biogeographical regions based on the species they harbour. This thesis also estimates butterfly sampling effort and associated biases, evaluate the national coverage of the network of protected areas for these insects, and develop initial models of potential distribution for rare or threatened species. Enrique also co-directed, together with Ana Luz Márquez and Raimundo Real from the Universidad de Málaga (<abbrev xlink:title="Universidad de Málaga">UMA</abbrev>), the thesis defended in 2022 by Antonio Pulido entitled “Application of fuzzy logic to the analysis of the spatial structure of diurnal butterfly populations in the Iberian Peninsula”. As a field entomologist, his tenacity was noteworthy. To find the elusive larvae of the satyrids he studied, he would amble through clumps of grass with an enormous comb and collect the caterpillars in a metal dish. His tremendous effort to complete the butterfly data for Spanish areas lacking records was also remarkable. He took advantage of both personal and university trips to stop over in unexplored regions. The butterfly database used for the two editions of the “Atlas de Mariposas Españolas” compiles his numerous field observations, scattered throughout Spain. With endless creativity, Enrique got surprising and often brilliant proposals and initiatives. He made excellent sketches and possessed a capacity for abstraction and synthesis far superior to most of us mere mortals, making him such a pleasure to work with. He constantly surprised us with ingenious ideas, collages, humorous posters, and those small details that always brought a smile to our faces. He was not one to delegate, but he loved helping his colleagues and family. His passion was research, without neglecting his teaching; his classes were engaging and distinctive, designed to cultivate critical thinking skills in students who questioned everything. As a colleague, Enrique García-Barros was an exceptional person. He always exuded an optimism, tinged with skepticism and irony. His idiosyncrasies included his humming and his soft whistling in the department corridors which were legendary, as was his intelligent humour with colleagues and students. It was a pleasure to work with him; he was always willing, attentive, a good listener, and generous. Discreet and modest, even somewhat introverted, he rarely expressed his worries, which, like anyone else, he surely had. He faced his short and fatal illness with the same tranquillity as the rest of his life. His family, friends, colleagues, and all who knew him will miss him utterly. Rest in peace, Enrique. <bold>Y ME DECÍS LUEGO</bold> Se marcha Enrique, El pasillo queda silente, Sin el silbo, sin el canturreo, Quizá un lejano rumor de voces inconcretas. Luego Se apaga la luz y todos lloran, Pero enseguida se encienden candiles y bengalas; Son chispas, fogonazos, flashes, El fondo sigue siendo oscuro y silencioso Pero hay luces fugaces Que alegran las miradas, Ecos lejanos, Pero perceptibles, Que recuerdan que no se fue del todo, Que permanece tanto En livianos garabatos, Como en dibujos olvidados Y, cómo no, en sus escritos, Sólidos, magníficos, creativos, De portentosa imaginación Y geniales ocurrencias. No, no se fue del todo, Escuchad esos ecos, Recordad esas vivencias Y me decís luego. <bold>AND TELL ME LATER</bold> Enrique leaves, The hallway falls silent, Without the whistling, without the humming, Perhaps a distant murmur of indistinct voices. Then The light goes out and everyone cries, But soon candles and flares are lit; They are sparks, flashes, bursts, The background remains dark and silent But there are fleeting lights That brighten our gazes, Distant echoes, But perceptible, That remind us he didn’t leave completely, That he remains both In light scribbles, As in forgotten drawings And, of course, in his writings, Solid, magnificent, creative, Of prodigious imagination And brilliant insights. No, he didn’t leave completely, Listen to those echoes, Remember those experiences And tell me later. </sec> </body> <back> <app-group> <app id="app1"> <title>Appendix 1. List of publications

Calle J, García-Barros E (1981) Nuevas adiciones a los Noctuidae de Guadalajara (contribución a los Noctuidae españoles n° 31). SHILAP Revista de lepidopterología 9(34): 124.

García-Barros E (1981) Citas nuevas o de interés para la provincia de Guadalajara. SHILAP Revista de Lepidopterología 9(36): 289–290.

García-Barros E (1982a) Las mariposas de la Alcarria. Tesina de licenciatura inédita. Universidad Autónoma de Madrid. Madrid. 288 pp.

García-Barros E (1982b) Ginopigios de los Papiliónidos (Papilionidae Latreille, 1809) Ibéricos. SHILAP Revista de Lepidopterología 10(38): 137–140.

García-Barros E (1982c) Hespéridos de la región de la Alcarria. SHILAP Revista de Lepidopterología 10(40): 287–291.

García-Barros E (1982d) Experiencia de cría del LasiocampidaePachygastria trifolii (D & Schiff, 1775) en regímenes de temperatura artificiales. SHILAP Revista de lepidopterología 10(37): 59–62.

García-Barros E (1982e) Diversidad estacional en una taxocenosis de lepidópteros. SHILAP Revista de lepidopterología 10(39): 227–230.

Baixeras J, García-Barros E, González P (1983) Meteorus abscissus Thomson, 1895 (Hym., Braconidae), primera cita para España. Pp.: 65–68. . In: Universidad de León (Eds), Actas I Congreso Ibérico de Entomología. Servicio de publicaciones de la Universidad de León, León.

García-Barros E (1984a) Ropalóceros (Lep. Papilionoidea) de la Alcarria (Guadalajara): fenología y abundancia. Boletín de la Asociación española de Entomología 8: 143–149.

García-Barros E (1984b) Morfología de las fases preimaginales y observaciones sobre la biología de Oxicesta serratae Zerny, 1927 (Lep., Noctuidae). Boletín de la Asociación española de Entomología 8: 111–120.

García-Barros E (1985a) Identificación de la larva y datos biológicos de Eilema uniola Rambur, 1858. Caracterización provisional de la larva del género Eilema Hübner (Lep., Arctiidae). Boletín de la Asociación española de Entomología 9: 223–237.

García-Barros E (1985b) Determinación de la larva de Cryphia muralis (Forst., 1771) (Lepidoptera, Noctuidae). Actas II Congreso Ibérico de Entomología. Boletim Sociedade Portuguesa de Entomologia 1(suppl. 1): 351–358.

García-Barros E (1986a) Notas sobre el comportamiento de las hembras de Brintesia circe (Fabricius) (Lep., Nymphalidae-Satyrinae): estridulación y agresividad. Actas VIII Jornadas de la Asociación española de Entomología, Sevilla: 849–854.

García-Barros E (1986b) Morfología externa de las pupas de Hipparchia Fabricius y Brintesia Frühstorfer (Lep., Satyridae, Satyrinae). Boletín de la Asociación española de Entomología 10: 339–353.

García-Barros E (1987a) Morphology and chaetotaxy of the 1st instar larvae of 6 species of the Satyrus (sl) series (Lepidoptera, Nymphalidae). Systematic Entomology 12(3): 335–344.

García-Barros E (1987b) Estudio comparativo de la biología y morfología de cuatro especies de Hipparchia Fabricius (Lepidoptera, Satyridae). Tesis doctoral, edición en microficha. Facultad de Ciencias, Universidad Autónoma de Madrid, 379 pp.

García-Barros E (1987c) Observaciones sobre la biología de Maniola jurtina (L., 1758) en el centro de la Península Ibérica: fenología general del ciclo biológico, duración del periodo de prepuesta, y fecundidad potencial de las hembras. Boletín de la Asociación española de Entomología 11: 235–247.

García-Barros E (1987d) Descripción de las fases preimaginales y biología de Paidia murina (Hb., 1822) (Lepidoptera, Arctiidae). Graellsia 42: 175–187.

García-Barros E (1988) Delayed ovarian maturation in the butterfly Hipparchia semele as a possible response to summer drought. Ecological Entomology 13(4): 391–398.

García-Barros E (1989a) Estudio comparativo de los caracteres biológicos de dos satirinos, Hipparchia statilinus (Hufnagel, 1766) e H. semele (L., 1758) (Lepidoptera, Nymphalidae, Satyrinae). Miscel·lània Zoologica 13: 85–96.

García-Barros E (1989b) Morfología larvaria de Hipparchia (Pseudotergumia) fidia (L., 1767) (Lepidoptera, Nymphalidae). Nouvelle Revue d’Entomologie (N.S.), 6(1): 71–83.

García-Barros E (1989c) Biología de Hipparchia (Pseudotergumia) fidia (L., 1767) (Lepidoptera: Nymphalidae, Satyrinae). SHILAP Revista de lepidopterología 17(66): 223–234.

García-Barros E (1989d) Phenological synchronization and adaptation in five satyrine butterflies from Central Spain. Nota Lepidopterologica 12(suppl. 1): 67.

García-Barros E, Viejo JL (1989) An attempt at the classification of six species of Satyrus (s.l.) based on morphological characters of the early stages. Nota Lepidopterologica 12(suppl. 1): 66.

Ibero C, García-Barros E, Yela JL (1989) Recopilación de la información y propuesta para un estudio de la distribución de las especies de papilionoideos y hesperioideos en la provincia de Guadalajara. SHILAP Revista de Lepidopterología 17(65): 61–70.

Viejo JL, Martín J, García-Barros E, Munguira ML (1990) The status of the Iberian Lepidoptera listed in the appendixes to the Bern Convention. Pp.: 42–45. . In: Council of Europe (Eds), Colloquy on the Berne Convention invertebrates and their conservation. Environmental encounters series n° 10. Council of Europe Publishing, Strasbourg.

García-Barros E, Martin J (1991) Immature stages of Hipparchia Fabricius and the systematics of the Satyrus series (Lepidoptera, Nymphalidae, Satyrinae). Systematic Entomology 16(4): 407–426.

Viejo JL, Martín J, García-Barros E, Munguira ML (1991) The Bern Convention and the Spanish Lepidoptera. Pp.: 428–429. . In: N. M. Collins & J. A. Thomas (Eds), The Conservation of Insects and their Habitats. Academic Press, Londres (Reino Unido).

García-Barros E (1992a) Evidence for geographic variation of egg size and fecundity in a Satyrine butterfly, Hipparchia semele (L.) (Lepidoptera, Nymphalidae-Satyrinae). Graellsia 48: 45–52.

García-Barros E (1992b) On egg size and fecundity of Satyrid butterflies. Pp.: 280–285. In: T. Pavlicek-van Beek, A. H. Ovaa & J. G. Van der Made (Eds), Future of butterflies in Europe: Strategies for survival. Agricultural University, Publication Serv., Wageningen (Holanda).

García-Barros E (1992c) Fases preimaginales, distribución y ciclo vital de Coscinia romeii Sagarra, 1924 (Lepidoptera, Arctiidae). EOS 68(2): 137–145.

Viejo JL, Martin J, García-Barros E, Munguira ML (1992) Diversidad de mariposas en el Parque Regional del Manzanares (Madrid) (Lepidoptera: Papilionoidea & Hesperioidea). Boletim Sociedade Portuguesa de Entomologia supl. 3(1): 201–212.

García-Barros E, Martín J, Munguira ML, Viejo JL (1993a) Fauna de mariposas y macroheteróceros del Parque Regional de la Cuenca Alta del Manzanares, Madrid, España. SHILAP Revista de Lepidopterología 21(82): 119–129.

García-Barros E, Munguira ML, Martín J, Viejo JL (1993b) Maculinea nausithous (Bergsträsser, 1779) en Madrid. SHILAP Revista de Lepidopterología 21(84): 255–256.

Nylin S, Wiklund C, Wickman PO, García-Barros E (1993) Absence of trade-offs between sexual size dimorphism and early male emergence in a butterfly. Ecology 74(5): 1414–1427.

García-Barros E (1994) Egg size variation in European Satyrine butterflies (Nymphalidae, Satyrinae). Biological Journal of the Linnean Society 51(3): 309–324.

García-Barros E, Martín J (1994) Morfología de los tubérculos mesotorácicos de las pupas de algunos lepidópteros (Lepidoptera: Nymphalidae, Hesperiidae, Sphingidae). SHILAP Revista de lepidopterología 22(87): 249–258.

Fernández-Rubio F, García-Barros E (1995) Taxonomic situation of the species of the genus Pararge Hübner, 1819 - P. aegeria, P. xiphia, and P. xiphioides - (Lepidoptera: Satyridae) in the Macaronesian Islands. Boletim do Museo Municipal do Funchal 47(260): 39–50.

García-Barros E, Martin J (1995) The eggs of european satyrine butterflies (Nymphalidae): external morphology and its use in systematics. Zoological Journal of the Linnean Society 115(1): 73–115.

Martín Cano J, Ferrín JM, García-Barros E, García-Ocejo A, Gurrea P, Luciáñez MJ, Munguira ML, Pérez Barroeta F, Ruiz Ortega M, Sanz Benito MJ, Simón JC, Viejo JL (1995) Las comunidades de insectos del Parque Regional de la Cuenca Alta del Manzanares (centro de España): estado de conservación. Graellsia 51: 101–111.

Wickman PO, García-Barros E, Rappegeorge C (1995) The location of landmark leks in the small heath butterfly, Coenonympha pamphilus evidence against the hot-spot model. Behavioral Ecology 6(1): 39–45.

Alonso-Zarazaga MA, Compte A, García-Barros E, Gurrea P, Izquierdo I, López M, Luciáñez MJ, Llorente V, Martín J, Martín MP, Martín F, Mingo E, Nieves JL, Rey C, Sanz MJ, Viejo JL (1996) Avances en entomología ibérica. Asociación Española de Entomología. Sociedade Portuguesa de Entomologia. Actas VI Congreso Ibérico de Entomología. Madrid. 502 pp.

García-Barros E (1996) Spot distribution in Maniola jurtina (Linnaeus, 1758) (Lepidoptera: Nymphalidae, Satyrinae) in Central Spain. Nota Lepidopterologica 19: 152–164.

García-Barros E, Munguira ML (1997) Uncertain branch lengths, taxonomic sampling error, and the egg to body size allometry in temperate butterflies (Lepidoptera). Biological Journal of the Linnean Society 61(2): 201–221.

Munguira ML, García-Barros E, Martín J (1997a) Plantas nutricias de los licénidos y satirinos españoles (Lepidoptera: Lycaenidae y Nymphalidae). Boletín de la Asociación española de Entomología 21(1–2): 29–53.

Munguira ML, Martín J, García-Barros E, Viejo JL (1997b) Use of space and resources in a Mediterranean population of the butterfly Euphydryas aurinia. Acta Oecologica 18(5): 597–612.

García-Barros E, Martín J, Munguira ML, Viejo JL (1998) Relación entre espacios protegidos y la diversidad de la fauna de mariposas (Lepidoptera: Papilionoidea et Hesperioidea) en la comunidad de Madrid: Una evaluación. Ecología 12: 423–439.

García-Barros E (1999a) Implicaciones ecológicas y evolutivas del tamaño en los artrópodos. Boletín de la Sociedad Entomológica Aragonesa 26: 657–678.

García-Barros E (1999b) Filogenia y evolución de Lepidoptera. Boletín de la Sociedad Entomológica Aragonesa 26: 475–483.

García-Barros E, Munguira ML (1999) Faunística de mariposas diurnas en España peninsular. Áreas poco estudiadas: una evaluación en el umbral del Siglo XXI (Lepidoptera: Papilionoidea & Hesperiidae). SHILAP Revista de lepidopterología 27(106): 189–202.

Garcia-Pereira P, García-Barros E, Munguira M (1999) Evaluación del conocimiento de la fauna de mariposas de Portugal continental. SHILAP Revista de lepidopterología 27(106): 225–231.

Munguira ML, García-Barros E (1999) National compilers. In: C.A.M. van Swaay & M.S. Warren (Eds), Red Data Book of European Butterflies (Rhopalocera). Nature and environment, No. 99. Council of Europe, Strasbourg.

García-Barros E (2000a) Body size, egg size, and their interspecific relationships with ecological and life history traits in butterflies (Lepidoptera: Papilionoidea, Hesperioidea). Biological Journal of the Linnean Society 70(2): 251–284.

García-Barros E (2000b) Comparative data on the adult biology, ecology and behaviour of species belonging to the genera Hipparchia, Chazara and Kanetisa in central Spain (Nymphalidae: Satyrinae). Nota Lepidopterologica 23(2): 119–140.

García-Barros E (2000c) Notas sobre la biología de los adultos de Pandoriana pandora (Dennis & Schiffermüller, 1775) en la España central (Lepidoptera: Nymphalidae). SHILAP Revista de Lepidopterología 28(109): 97–102.

García-Barros E (2000d) Egg size in butterflies (Lepidoptera: Papilionoidea and Hesperiidae): a summary of data. Journal of Research on the Lepidoptera 35: 90–136.

García-Barros E (2000e) Clima y tamaño en mariposas diurnas (Lepidoptera: Papilionoidea). Boletín de la Asociación española de Entomología 24: 47–64.

García-Barros E, (coord) (2000f) Prácticas de Zoología General. Guiones de prácticas. Documentos de trabajo n° 25. Ediciones de la Universidad Autónoma de Madrid. Madrid. 191 pp.

García-Barros E, Garcia-Pereira P, Munguira ML (2000) The geographic distribution and state of butterfly faunistic studies in Iberia (LepidopteraPapilionoideaHesperioidea). Belgian Journal of Entomology 2: 111–124.

Martín J, García-Barros E, Gurrea P, Luciáñez MJ, Munguira M, Sanz MJ, Simón JC (2000) High endemism areas in the Iberian Peninsula. Belgian Journal of Entomology 2: 47–57.

Munguira ML, Martín J, García-Barros E (2000a) Maculinea nausithous (Bergsträsser, 1779). Pp.: 67–71. In: E. Galante & J. R. Verdú (Eds), Los artrópodos de la “Directiva Hábitat” en España. Organismo Autónomo de Parques Nacionales, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2000b) Plebicula golgus (Hübner, 1813). Pp.: 72–75. In: E. Galante & J. R. Verdú (Eds), Los artrópodos de la “Directiva Hábitat” en España. Organismo Autónomo de Parques Nacionales, Ministerio de Medio Ambiente, Madrid.

García-Barros E, Herranz J (2001) Nuevas localidades de Proserpinus proserpina (Pallas, 1772) y Graellsia isabelae (Graells, 1849). SHILAP Revista de Lepidopterología 29(114): 183–184.

Garcia-Pereira P, Maravalhas E, García-Barros E (2001) Dos lepidópteros nuevos para Portugal: Adscita (Adscita) bolivari (Ajenjo, 1937) (Zygaenidae) y Brenthis ino (Rottemburg, 1775) (Nymphalidae). Zapateri Revista aragonesa de entomología 9: 115–116.

Munguira ML, Martín J, Orueta D, Viejo JL, García-Barros E (2001) Maculinea nausithous (Bergstrasser, 1779). Pp.: 163–173. In: M. Ramos, D. Bragado & J. Fernandez (Eds), Los invertebrados no insectos de la “Directiva Hábitat” en España. Organismo Autónomo de Parques Nacionales, Ministerio de Medio Ambiente, Madrid.

García-Barros E (2002) Taxonomic patterns in the egg to body size allometry of butterflies and skippers (Papilionoidea & Hesperiidae). Nota Lepidopterologica 25(2/3): 161–175.

García-Barros E, Gurrea P, Luciáñez MJ, Martín Cano J, Munguira ML, Moreno JC, Sainz H, Sanz MJ, Simón JC (2002) Parsimony analysis of endemicity and its application to animal and plant geographical distributions in the Ibero-Balearic region (western Mediterranean). Journal of Biogeography 29(1): 109–124.

García-Barros E (2003) Mariposas diurnas endémicas de la región paleártica occidental: Patrones de distribución y su análisis mediante parsimonia (Lepidoptera, Papilionoidea). Graellsia 59(2–3): 233–258.

Garcia-Pereira P, García-Barros E (2003) A evolução das borboletas. Pp.: 28–34. In: E. Maravalhas (Eds), As borboletas de Portugal. Apollo Books, Stenstrup (Dinamarca).

Garcia-Pereira P, García-Barros E, Maravalhas E (2003a) Actualización de la distribución geográfica de los Zygaenidae en Portugal continental (Lepidoptera: Zygaenidae). SHILAP Revista de Lepidopterología 31(122): 157–168.

Garcia-Pereira P, García-Barros E, Munguira ML (2003b) Patrones de distribución de las mariposas diurnas en Portugal (Lepidoptera, Papilionoidea, Hesperiidae, Zygaenidae). Graellsia 59: 259–271.

Munguira ML, García-Barros E, Martín J (2003) Prime butterfly areas in Spain. Pp.: 531–554. In: C.A.M. van Swaay & M.S. Warren (Eds), Prime butterfly areas in Europe: Priority sites for conservation. National Reference Centre for Agriculture, Nature and Fisheries. Ministry of agriculture, Nature management and Fisheries, The Netherlands.

Romo H, García-Barros E, Chaves P, Garcia-Pereira P, Maravalhas E (2003) Distribución actualizada de las especies de Coenonympha Hübner, [1819] y Melanargia Meigen, [1828] en la Península Ibérica (Lepidoptera: Nymphalidae, Satyrinae). SHILAP Revista de lepidopterología 31(124): 329–354.

García-Barros E (2004) Implicaciones ecológicas y evolutivas del tamaño en los artrópodos. Pp.: 203 - 228. In: J.E. Llorente Bousquets, J.J. Morrone, O. Yáñez Ordóñez & I. Vargas Fernández (Eds), Biodiversidad, taxonomía y biogeografía de artrópodos: Hacia una síntesis de su conocimiento. Vol. IV. Facultad de Ciencias, Universidad Nacional Autónoma de México, México, D.F.

García-Barros E, Chaves P, Coles S, Wright L (2004a) Distribución ibérica de once especies de satirinos (Lepidoptera, Nymphalidae, Satyrinae). SHILAP Revista de lepidopterología 32(125): 57–79.

García-Barros E, Munguira ML, Martín J, Romo H, Garcia-Pereira P, Maravalhas ES (2004b) Atlas de las mariposas diurnas de la Península Ibérica e islas Baleares (Lepidoptera: Papilionoidea & Hesperioidea). Atlas of the butterflies of the Iberian Peninsula and Balearic Islands (Lepidoptera: Papilionoidea & Hesperioidea). Monografias de la SEA, vol 11. Sociedad Entomológica Aragonesa. Zaragoza. 228 pp.

Hortal J, Garcia-Pereira P, García-Barros E (2004) Butterfly species richness in mainland Portugal: predictive models of geographic distribution patterns. Ecography 27(1): 68–82.

Maravalhas E, Pires P, Garcia-Pereira P, Romão F, García-Barros E (2004) Ampliação do conhecimento dos ropalóceros (Lepidoptera: Hesperioidea e Papilionoidea) do nordeste de Portugal. Boletín de la Sociedad Entomológica Aragonesa 35: 137–142.

García-Barros E (2005) Size-dependent, continuous response of larval growth rates to photoperiod in the Satyrine Nymphalid Coenonympha pamphilus (L., 1758). Pp.: 39–40. In: E. Kuehn, R. Feldmann, J. Thomas & J. Settele (Eds), Studies on the ecology and conservation of butterflies in Europe. 1: General concepts and case studies. Pensoft, Sofia-Moscow.

García-Barros E, Romo H (2005) Latitude, longitude, and the evolution of Iberian butterfly faunistics (Lepidoptera). A preliminary test for shifts in distribution areas in the Western Mediterranean. Pp.: 52–54. In: E. Kuehn, R. Feldmann, J. Thomas & J. Settele (Eds), Studies on the ecology and conservation of butterflies in Europe. 1: General concepts and case studies. Pensoft, Sofia-Moscow.

Romo H, García-Barros E (2005) Distribución e intensidad de los estudios faunísticos sobre mariposas diurnas en la Península Ibérica e islas Baleares (Lepidoptera, Papilionoidea y Hesperioidea). Graellsia 61(1): 37–50.

Romo H, García-Barros E, Lobo JM (2005) What areas to protect: biases in the historic record of Iberian butterflies faunistics. Pp.: 25–26. In: E. Kuehn, R. Feldmann, J. Thomas & J. Settele (Eds), Studies on the ecology and conservation of butterflies in Europe. 1: General concepts and case studies. Pensoft, Sofia-Moscow.

Simón Benito JC, Espantaleón D, García-Barros E (2005) Stachorutes cabagnerensis n. sp., Collembola (Neanuridae) from Central Spain, and a preliminary approach to phylogeny of genus. Animal Biodiversity and Conservation 28(2): 149–157.

García-Barros E (2006a) Number of larval instars and sex-specific plasticity in the development of the small heath butterfly, Coenonympha pamphilus (Lepidoptera: Nymphalidae). European Journal of Entomology 103(1): 47–53.

García-Barros E (2006b) Within and between species scaling in the weight, water, carbon and nitrogen contents of eggs and neonate larvae of twelve satyrine butterflies (Lepidoptera: Nymphalidae). European Journal of Entomology 103(3): 559–568.

García-Barros E, Martín J, Munguira ML (2006a) Lopinga achine (Scopoli, 1763). Pp.: 243–244. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

García-Barros E, Martín J, Munguira ML (2006b) Coscinia romeii Sagarra, 1924. Pp.: 220–221. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Martín J, Munguira ML, García-Barros E (2006a) Pyrgus cinarae (Rambur, 1840). Pp.: 222. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Martín J, Munguira ML, García-Barros E (2006b) Pyrgus sidae (Esper, 1782). Pp.: 223. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006a) Maculinea nausithous (Bergsträsser, 1779). Pp.: 227–228. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006b) Agriades zullichi (Hemming, 1933). Pp.: 224–225. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006c) Chazara prieuri (Pierret, 1837). Pp.: 242. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006d) Lycaena helle (Dennis y Schiffermüller, 1775). Pp.: 226. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006e) Maculinea rebeli (Hirsche, 1904). Pp.: 229. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006f) Plebejus hespericus (Rambur, 1840). Pp.: 230–232. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín J, García-Barros E (2006g) Polyommatus golgus (Hübner, 1813). Pp.: 233–234. In: E. Galante & J. R. Verdú (Eds), Libro Rojo de los Invertebrados de España. Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Romo H, García-Barros E, Lobo JM (2006a) Identifying recorder-induced geographic bias in an Iberian butterfly database. Ecography 29(6): 873–885.

Romo H, García-Barros E, Munguira M (2006b) Distribución potencial de trece especies de mariposas diurnas amenazadas o raras en el área ibero-balear (Lepidoptera: Papilionoidea & Hesperioidea). Boletín de la Asociación española de Entomología 30(3–4): 25–49.

Romo H, Munguira M, García-Barros E (2007) Area selection for the conservation of butterflies in the Iberian Peninsula and Balearic Islands. Animal Biodiversity and Conservation 30.1: 7–27.

García-Barros E, Fartmann T (2009) Butterfly oviposition: sites, behaviour and modes. Pp.: 29–42. In: J. Settele, T. Shreeve, M. Konvička & H. Van Dyck (Eds), Ecology of Butterflies in Europe. Cambridge University Press, Cambridge.

Martín Cano J, García-Barros E, Munguira ML (2009) Lycaena helle (Dennis y Schiffermüller, 1775). Pp.: 180–184. In: J. R. Verdú & E. Galante (Eds), Atlas de los Invertebrados Amenazados de España (Especies En Peligro Crítico y En Peligro). Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Modolell J, Munguira ML, García-Barros E (2009) Presence and phenotypic variability of Palmaris gustavi (Staudinger, 1898) in northern Chile, with notes on Palmaris penai (Hayward, 1967) (Lepidoptera, Nymphalidae, Satyrinae). Boletín de la Sociedad Entomológica Aragonesa 44: 351–356.

Munguira ML, García-Barros E, Martín Cano J (2009a) Butterfly herbivory and larval ecology. Pp.: 43 - 54. In: J. Settele, T. Shreeve, M. Konvička & H. Van Dyck (Eds), Ecology of Butterflies in Europe. Cambridge University Press, Cambridge.

Munguira ML, Martín Cano J, García-Barros E, López Pajarón J (2009b) Polyommatus golgus (Hübner, [1813]). Pp.: 185–189. In: J. R. Verdú & E. Galante (Eds), Atlas de los Invertebrados Amenazados de España (Especies En Peligro Crítico y En Peligro). Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

Munguira ML, Martín Cano J, García-Barros E, López Pajarón J (2009c) Agriades zullichi Hemming, 1930. Pp.: 176–179. In: J. R. Verdú & E. Galante (Eds), Atlas de los Invertebrados Amenazados de España (Especies En Peligro Crítico y En Peligro). Dirección General para la Biodiversidad, Ministerio de Medio Ambiente, Madrid.

García-Barros E, Romo H (2010) The relationship between geographic range size and life history traits: is biogeographic history uncovered? A test using the Iberian butterflies. Ecography 33(2): 392–401.

Palacios Vargas JG, García-Barros E, Simon Benito JC (2010) Phylogeny of the genus Palmanura (Collembola: Neanuridae). Cladistics 26(5): 482–496.

Romo H, García-Barros E (2010) Biogeographic regions of the Iberian Peninsula: butterflies as biogeographical indicators. Journal of Zoology 282(3): 180–190.

García-Barros E, Munguira ML, Martín Cano J, López Pajarón J (2011a) Coscinia romeii Sagarra, 1924. Pp.: 458–463. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados Amenazados de España (Especies Vulnerables), volumen I. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio rural y Marino, Madrid.

García-Barros E, Murria E, Romo H, Munguira ML, Martín J (2011b) Chazara prieuri (Pierret, 1837). Pp.: 1274–1281. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados amenazados de España (Especies Vulnerables), volumen II. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio Rural y Marino, Madrid.

García-Barros E, Pascual J, Romo H (2011c) Contribución corológica de un recurso de la Red: Biodiversidad Virtual y las mariposas diurnas íbero-baleares (Lepidoptera, Papilionoidea). Boletín de la Asociación española de Entomología 35(3–4): 457–475.

Mortera H, García-Barros E, Romo H, Munguira ML, Martín Cano J, Lamata F (2011) Lopinga achine (Scopoli, 1763). Pp.: 478–483. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados Amenazados de España (Especies Vulnerables), volumen I. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio rural y Marino, Madrid.

Munguira ML, González J, Romo H, Martín J, García-Barros E (2011a) Phengaris alcon (Denis y Schiffermüller, 1775). Pp.: 1248–1257. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados amenazados de España (Especies Vulnerables), volumen II. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio Rural y Marino, Madrid.

Munguira ML, Romo H, Martín J, García-Barros E (2011b) Phengaris nausithous (Bergsträsser, 1779). Pp.: 1258–1264. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados amenazados de España (Especies Vulnerables), volumen II. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio Rural y Marino, Madrid.

Munguira ML, Romo H, Pérez R, Martín Cano J, García-Barros E (2011c) Plebejus hespericus (Rambur, 1840). Pp.: 1265–1273. In: J. R. Verdú, C. Numa & E. Galante (Eds), Atlas y Libro Rojo de los Invertebrados Amenazados de España (Especies Vulnerables), volumen II. Dirección General de Medio Natural y Política Forestal, Ministerio de Medio Ambiente, Medio rural y Marino, Madrid.

García-Barros E, Meneguz M (2012) Structure of the wing ventral scales of Coenonympha Hubner, [1819] with special reference to C. pamphilus (L., 1758) and its morphotype lyllus (Esper, 1805) (Lepidoptera: Nymphalidae). SHILAP-Revista De Lepidopterologia 40(159): 279–293.

García-Barros E, Romo H, Molina M (2012) Relationship between geographic rarity and perception of threat in Iberian butterflies. Journal of Insect Conservation 16(3): 355–366.

Real R, Márquez AL, Romo H, Moreno JC, Acevedo P, García-Barros E (2012) Fuzzy logic as a key conceptual tool for modelling the distribution of interacting species in climate change scenarios. Conference program and abstracts International Biogeography Society 6th Biennial Meeting Miami. Frontiers of biogeography 4 (issue 5, suppl.1): 60.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012a) Maculinea nausithous. Pp.: 1–49. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012b) Plebicula golgus. Pp.: 1–47. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012c) Parnassius mnemosyne. Pp.: 1–47. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012d) Proserpinus proserpina. Pp.: 1–45. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012e) Parnassius apollo. Pp.: 1–65. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012f) Maculinea arion. Pp.: 1–55. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012g) Lopinga achine. Pp.: 1–45. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012h) Euphydryas aurinia. Pp.: 1–67. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012i) Eriogaster catax. Pp.: 1–45. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

Romo H, García-Barros E, Martín J, Ylla J, Munguira ML (2012j) Graellsia isabelae. Pp.: 1–53. In: VV.AA. (Eds), Bases ecológicas preliminares para la conservación de las especies de interés comunitario en España: Invertebrados. Ministerio de Medio Ambiente, y Medio Rural y Marino, Madrid.

García-Barros E, Munguira ML, Stefanescu C, Vives A (2013) Lepidoptera: Papilionoidea. Pp.: 1213. In: M. A. Ramos, J. Alba, X. Bellés, J. Gosálbez, A. Guerra, E. Macpherson, J. Serrano & J. Templado (Eds), Fauna Iberica, 37, Museo Nacional de Ciencias Naturales, CSIC, Madrid.

Romo H, Sanabria P, García-Barros E (2013) Predicting climate change impacts on Lepidoptera species distribution. The case of genus Boloria Moore, 1900 in the Iberian Peninsula (Lepidoptera: Nymphalidae). SHILAP-Revista de lepidopterologia 41(162): 267–286.

Cancela JP, Munguira ML, García-Barros E (2014) Immature stages and adult morphology as a contribution to the phylogeny of the West Palaearctic lycaenid butterflies (Lepidoptera: Lycaenidae). Annales de la Societe Entomologique de France 50(3–4): 418–430.

Romo H, Camero-R, E, García-Barros E, Munguira ML, Martin Cano J (2014a) Recorded and potential distributions on the Iberian Peninsula of species of Lepidoptera listed in the Habitats Directive. European Journal of Entomology 111(3): 407–415.

Romo H, García-Barros E, Márquez AL, Moreno JC, Real R (2014b) Effects of climate change on the distribution of ecologically interacting species: butterflies and their main food plants in Spain. Ecography 37(11): 1063–1072.

Dincă V, Backström N, Dapporto L, Friberg M, García-Barros E, Hebert PDN, Hernández-Roldán J, Hornett E, Lukhtanov V, Marec F, Montagud S, Munguira ML, Olofsson M, Šíchová J, Talavera G, Vicente-Arranz JC, Vila R, Wiklund C (2015) DNA barcodes highlight unique research models in European butterflies. Genome 58(5): 163–303.

Dincă V, Montagud S, Talavera G, Hernández-Roldán J, Munguira ML, García-Barros E, Hebert PDN, Vila R (2015) DNA barcode reference library for Iberian butterflies enables a continental-scale preview of potential cryptic diversity. Scientific Reports 5(1): 12395.

García-Barros E (2015) Multivariate indices as estimates of dry body weight for comparative study of body size in Lepidoptera. Nota Lepidopterologica 38(1): 59–74.

García-Barros E, Romo H, Sarto I, Monteys V, Munguira ML, Baixeras J, Vives Moreno A, Yela García JL (2015) Clase Insecta: Orden Lepidoptera. Revista IDE@ (Ibero Diversidad Entomológica @ccesible)- SEA, 65: 1–21.

Munguira ML, Martin J, García-Barros E, Shahbazian G, Pablo Cancela J (2015) Morphology and morphometry of Lycaenid eggs (Lepidoptera: Lycaenidae). Zootaxa 3937(2): 201–247.

Slancarova J, Garcia-Pereira P, Fric ZF, Romo H, García-Barros E (2015) Butterflies in Portuguese ‘montados’: relationships between climate, land use and life-history traits. Journal of Insect Conservation 19(5): 823–836.

García-Barros E (coord.) (2016) Prácticas de Zoología General. Guiones de prácticas. 2ª edición. Documentos de trabajo n° 101. Ediciones de la Universidad Autónoma de Madrid. Madrid, 182 pp.

Munguira ML, Barea-Azcón JM, Castro-Cobo S, García-Barros E, Miteva S, Olivares J, Romo H (2017) Ecology and recovery plans for the four Spanish endangered endemic butterfly species. Journal of Insect Conservation 21(3): 423–437.

Titeux N, Maes D, Van Daele T, Onkelinx T, Heikkinen RK, Romo H, García-Barros E, Munguira ML, Thuiller W, van Swaay CAM, Schweiger O, Settele J, Harpke A, Wiemers M, Brotons L, Luoto M (2017) The need for large-scale distribution data to estimate regional changes in species richness under future climate change. Diversity and Distributions 23(12): 1393–1407.

García-Barros E (2018) José Luis Viejo: Un Entomólogo en la UAM. Encuentros multidisciplinares 60 (extraordinario): 1–4.

Pulido-Pastor A, Luz Marquez A, García-Barros E, Real R (2018) Identification of potential source and sink areas for butterflies on the Iberian Peninsula. Insect Conservation and Diversity 11(5): 479–492.

Rosso A, Aragón P, Acevedo F, Doadrio I, García-Barros E, Lobo JM, Munguira ML, Monserrat VJ, Palomo J, Pleguezuelos JM, Romo H, Triviño V, Sánchez-Fernández D (2018) Effectiveness of the Natura 2000 network in protecting Iberian endemic fauna. Animal Conservation 21(3): 262–271.

Herrando S, Titeux N, Brotons L, Anton M, Ubach A, Villeros D, García-Barros E, Munguira ML, Godinho C, Stefanescu C (2019) Contrasting impacts of precipitation on Mediterranean birds and butterflies. Scientific Reports 9: 5680.

Hernández Hernández M, García-Barros E, Munguira ML, Mora Cabello de Alba A (2020) Estimación de la completitud en censos de mariposas diurnas con datos de seguimiento de BMS España (Lepidoptera, Papilionoidea). Boletín de la Asociación Española de Entomología 44(3–4): 451–470.

Hinojosa JC, Koubínová D, Dincă V, Hernández-Roldán J, Munguira ML, García-Barros E, Vila M, Alvarez N, Mutanen M, Vila R (2020) Rapid colour shift by reproductive character displacement in Cupido butterflies. Molecular Ecology 29(24): 4942–4955.

Middleton-Welling J, Dapporto L, García-Barros E, Wiemers M, Nowicki P, Plazio E, Bonelli S, Zaccagno M, Šašić M, Liparova J, Schweiger O, Harpke A, Musche M, Settele J, Schmucki R, Shreeve T (2020) A new comprehensive trait database of European and Maghreb butterflies, Papilionoidea. Scientific Data 7(1): 351.

Munguira ML, García-Barros E (2020) La red de Parques Nacionales como pieza clave del seguimiento en España. Boletín de la Red de Parques Nacionales 63: 11.

Mingarro M, Cancela JP, Burón-Ugarte A, García-Barros E, Munguira ML, Romo H, Wilson RJ (2021) Butterfly communities track climatic variation over space but not time in the Iberian Peninsula. Insect Conservation and Diversity 14(5): 647–660.

Pulido-Pastor A, Márquez AL, Guerrero JC, García-Barros E, Real R (2021) Metapopulation Patterns of Iberian Butterflies Revealed by Fuzzy Logic. Insects 12(5), 392.

Roth N, Hacker HH, Heidrich L, Friess N, García-Barros E, Habel JC, Thorn S, Müller J (2021) Host specificity and species colouration mediate the regional decline of nocturnal moths in central European forests. Ecography 44(6): 941–952.

García-Barros E, Cancela JP, Lobo JM, Munguira ML, Romo H (2022a) Forecasts of butterfly future richness change in the southwest Mediterranean. The role of sampling effort and non-climatic variables. Journal of Insect Conservation 26(4): 639–650.

García-Barros E, Viejo JL, Munguira ML (2022b) In memoriam: Fidel Fernandez-Rubio (13 Dec 1928–20 Apr 2022). Nota Lepidopterologica 45: 355–362.

Pla-Narbona C, Stefanescu C, Pino J, Cabrero-Sañudo FJ, García-Barros E, Munguira ML, Melero Y (2022) Butterfly biodiversity in the city is driven by the interaction of the urban landscape and species traits: a call for contextualised management. Landscape Ecology 37(1): 81–92.

Sánchez-Fernández D, Yela JL, Acosta R, Bonada N, García-Barros E, Guisande C, Heine J, Millán A, Munguira ML, Romo H, Zamora-Muñoz C, Lobo JM (2022) Are patterns of sampling effort and completeness of inventories congruent? A test using databases for five insect taxa in the Iberian Peninsula. Insect Conservation and Diversity 15(4): 406–415.

Viejo JL, García-Barros E, Munguira ML (2022) Necrológica Fidel Fernández Rubio (1928–2022). Boletín Real Sociedad Española Historia Natural 116: 243–249.

García-Barros E, Romo H, Castro-Cobo S, Ledesma E, Munguira ML, de Arce JI, Blázquez Caselles Á, Cancela JP, Cortizo Amaro C, Escobés Jiménez R, Farino T, García Carrillo A, Gómez Calmaestra R, González S, Hernández-Roldán JL, Martínez Pérez I, Miguel-Aristu J, Monasterio León Y, Montiel Pantoja C, Mora Cabello de Alba A, de Olano I, Ortiz Cervantes AS, Pérez Fernández R, Rodrigo Dapena J, Sánchez Á, Sanjurjo Franch MJ, Verovnik R, Vila Ujaldón R, Wilson RJ (2023) Actualización de la información sobre la distribución geográfica de las mariposas ibero-baleares (Lepidoptera, Papilionoidea): mapa de ignorancia. Boletín de la Sociedad Entomológica Aragonesa 72: 01–09.

Romo H, García-Barros E, Wilson RJ, Mateo RG, Munguira ML (2023) Modelling the scope to conserve an endemic-rich mountain butterfly taxon in a changing climate. Insect Conservation and Diversity 16(4): 451–467.

Álamo M, García-Barros E, Romo H (2024) Adult thermoregulatory behaviour does not provide, by itself, an adaptive explanation for the reflectance-climate relationship (Bogert’s pattern) in Iberian butterflies. Ecological Entomology 49(1): 77–90.

García-Barros E (2024) Travelling through Spain or Portugal next season? Lepidopterist-heroes are demanded for butterfly faunistics: fill the gap, or where to find the worst places to get poor -but highly prized- results! Newsletter of the Society for European Lepidopterology 69: 8–9.

Mañas-Jordá S, Acosta R, Ariño A, Baquero E, Bonada N, Galicia D, García-Barros E, García-Meseguer AJ, García-Roselló E, González M, Lobo JM, López-Rodríguez MJ, Martínez J, Millán A, Monserrat VJ, Munguira ML, Prieto CE, Romo H, Sánchez-Campaña C, Tierno de Figueroa JM, Yela JL, Sánchez-Fernández D (2025) IberArthro: A free online database compiling taxonomic and distributional data on Ibero-Balearic arthropods. Ecosistemas 34(2): 2955.

Sánchez-Fernández D, Mañas-Jordá S, García-Meseguer AJ, Acosta R, Ariño A, Baquero E, Bartomeus I, De Biurrun G, Bonada N, Galicia D, García-Barros E, García Paris M, García-Roselló E, Gil-Tapetado D, González M, Jurado Angulo P, López-Rodríguez MJ, Martínez J, Millán A, Monserrat VJ, Morano E, Munguira ML, Pallarés S, Pina F, Prieto C, Romo H, Sánchez-Campaña C, Sar P, Tierno De Figueroa JM, Yela JL, Lobo JM (2026) Priority areas for arthropod conservation in the Iberian Peninsula and Balearic Islands: Insights from a multi-taxon distributional database. Insect Conservation and Diversity 1–12. https://doi.org/10.1111/icad.70060

Marques V, Spilani L, Hinojosa J, Corbella C, Dincă V, Vodă R, Mesa Sánchez L, García-Barros E, Munguira ML, Talavera G, Dapporto L, Vila R(2026) The evolutionary retreats of Cyaniris semiargus - Beach or mountain? Contrasting paths across ecological gradients. Systematic Entomology 51: e70034. https://doi.org/10.1111/syen.70034

Munguira ML, Romo H, García-Barros E, Ledesma E, Castro S (coords.) (in press) Atlas y Libro Rojo de las Mariposas Españolas. MITECO. Madrid.

García-Meseguer AJ, Sánchez-Fernández D, Yela JL, García-Barros E, García-Roselló E, Mañas-Jordá S, Millán A, Munguira ML, Romo H, Lobo JM (submitted) Sampling bias masks cross-taxon congruence in Iberian insect species richness patterns. Biodiversity and Conservation.

De Gracia C, Ursul G, Cancela JP, García-Barros E, Munguira ML, Romo H, Wilson RJ, Mingarro M (submitted) Assessing butterfly habitat preferences and vegetation encroachment in Central Spain’s mountainous systems. Animal Biodiversity and Conservation.

Marques V, Spilania L, Hinojosa JC, Adams D, Dincă V, Dapporto L, Hernández-Roldán JL, Romo H, García-Barros E, Munguira ML, Vila M, Vila R (in prep.) False grayling, real diversity: genomic and morphological evidence for multiple species within Arethusana. Systematic Entomology.