Subject Editor: Sven Erlacher
Based on morphological and molecular analyses of “
Specimens were photographed before performing a standard method of dissection (
More than 300 specimens of
ZFMK Zoological Research Museum Alexander Koenig, Bonn
ZSM Bavarian State Collection of Zoology, Munich
PCDS Private collection Dirk Stadie, Lutherstadt Eisleben
PCR amplification and sequencing of 658 bp of COI mtDNA of the three freshly collected specimens of
Sequences were aligned using BOLD platform (
Lectotype (hereby designated in order to stabilize nomenclature) ♀, ”Syr. sept. [Turkey], Marasch, Achyr Dagh sept. Bertiz Jaila, 1800m, 09.-13.vi. [19]29, E. Pfeiffer leg.”, ”
Wingspan 25–29 mm, forewing length 13.2–15.0 mm; n=11. Apex pointed. Termen slightly rounded. Ground colour of forewing light ochre-brown to light grey-brown; basal and medial area darkened in half of all individuals, the others rather uniform; transverse lines well developed, distinct and dentate; postmedial line bordered distally with white scales; subterminal line narrow, whitish, more or less complete; terminal line fine dark brown, streak-like, interrupted at the veins; costa often suffused with whitish scales. Cell spots round, always present but often weak and hardly visible. Fringes slightly lighter than ground colour, chequered. Hindwing colour slightly lighter than forewing. Transverse lines usually indistinct except the postmedial and terminal lines. Cell spots usually absent. If present then very weak, developed as an elongate streak. Underside of both wings lighter than upper side, suffused with light ochre-brown scales. Transverse lines absent or strongly reduced except the postmedial and terminal lines. Cell spots on both wings weak, but always present. Head and frons unicolorously ochre-brown. Palpi reduced in size. Antennae of male slightly dentate in lateral view, those of female filiform. Tibia of forelegs without spurs, of mid-legs with one pair, of hindlegs with two pairs of spurs. Chaetosemata present.
Wing pattern.
Uncus flat, bifid, projections distally rounded. Valva broadly sclerotized at costa, with a rounded lobe and a deep, sub-apical incision. Apical projection thin, with small rounded tip. Juxta narrow and largely reduced, situated between the oval basal parts of the valvae and behind the pseudojuxta (only partly visible in Figs
Male and female genitalia.
Ductus bursae furrowed, slightly curved, near ostium remarkably widened, more sclerotized. Corpus bursae membranous (Fig.
Genetic similarity and interspecific distances are shown in the neighbour-joining tree (Fig.
Un-rooted neighbour-joining tree based on individuals belonging to six species of the genera
Interspecific distances between six species of the genera
1 | 2 | 3 | 4 | 5 | |
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9.3 | ||||
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2.4 | 9.1 |
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5. |
10.5 | 11.3 |
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10.7 | |
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4.2 | 8.6 |
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4.2 | 9.7 |
So far
Similar to other
Full-grown larva (L5) moderately slender, length 3 cm. Ground colour dorsally light green. Head beige. Epistigmatal line fine, whitish. Stigmatal line broad, ivory coloured, with a yellow tinge, indistinct. Stigmata bright yellow, bordered by a fine black margin. The whole body is covered scarcely with fine blackish setae, with small blackish patches at their bases. Ventrum uniform whitish-green (Fig.
We are grateful to Ralf Fiebig (Roßleben, Germany) for loan of his material and additional ecological information on this species. Our special thanks go to Paul Hebert (CCDB, University of Guelph, Canada) and his team for kindly and professionally performing sequencing of the material. Furthermore we thank Jaan Viidalepp (Tartu, Estonia) and Bernd Müller (Berlin, Germany) for competent help and additional information on the tribe
List of sequenced specimens, with identification, Sampling sites collecting data, Accession numbers, and process ID in BOLD database. Data taken from BOLD and generated by Axel Hausmann (1); Norbert Pöll (2); Petrányi Gergely (3), Dieter Stüning (4).
Taxon identification | Sampling site | Genbank Accession Nr. | Process ID (in BOLD database) |
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Italy, Calabria, Prov. Cosenza, Strada per Aieta, |
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Slovenia, Kras-Prešnica, |
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Greece, West-Macedonia, Vogatsikon (suedl. Kastoria), 29-May-2006, |
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Morocco, Province Souss-Massa-Draa, Agadir env.[ironments], 10km N[orth] Agadir, |
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Germany, Thuringia, S[üdlicher] Kyffhäuser, Ochsenburg, 08.V.2006, leg. et. coll. Dirk Stadie. |
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Italy, Calabria, Prov. Cosenza, Strada per Aieta, |
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Croatia, Primorje-Gorski Kotar, Krk Island, Bašćanska Draga, 12.VI.2000, leg. et. coll. Dirk Stadie. |
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Turkey, Dogu Anadolu, Erzincan, 20 km S. Erzincan, |
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Russia, Tuva, near Shagonar, Khairakan Mt., 565 m, |
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France.Corsica, Val d’Ese, 6km E Bastelica, 1600 m, 17.VII.2004, |
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Spain, Castilla-La Mancha, Guadalajara, 4km E Embid, 1075 m, 08.08.2007, leg. P. Skou |
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Spain, Aragon, Torres de Albarracin, |
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Spain, Aragon, Teruel, Sierra Albarracin, Moscardon, 1440 m, 22-May-2006, |
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Georgia, Kachetia, Tusheti, David Gazeta, 4.VII.2010, |
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Iran, Mazandaran, Resteh-Ye-Elborz, Mazandaran Pass, 2988 m, 22-Jun-2005, |
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Turkey, Dogu Anadolu, Province Malatya, Nemrut dagi northside, |
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Turkey, Dogu Anadolu, Province Malatya, Nemrut dagi northside, |
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Syria[Turkey], Achyr Dagh, Bertiz Jaila, 13.VII.1929, 1800 m, leg. E. Pfeiffer, Lectotype |
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Kyrgyzstan, Prov. Osh, Distr. Nookat, Kitschik-Alai, Abschyr-Say, 1820m, 20-Jun-2010, |
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Kyrgyzstan, Prov. Osh, Distr. Kara-Suu, Alai-Mountains, River Ak-Buura, Tarylga vic., 1535 m, 4-Jun-2010, |
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Uzbekistan, Tashkent, Parkent, reg. Surenata Mts., 900 m, 10-May-1995, |
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China, Beijing Shi, Yanqing, Dayushu, 520 m, 13-Jun-2007, |
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