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  • Article title
  • Abstract
  • Introduction
  • Material and methods
    • Study area
    • Study system: Erebia butterflies
    • Behavioural data
    • Statistical analyses
  • Results
    • Edge permeability
    • Time-of-day effects
  • Discussion
    • Road and species effects
    • Time effects
    • Conclusions
  • Acknowledgements
  • References
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